Japanese species of the minute litter bug genus Ceratocombus (Hemiptera, Heteroptera, Ceratocombidae), with descriptions of three new species

A taxonomic study of the Japanese Ceratocombus Signoret, 1852 was resumed for the first time in over 100 years, with descriptions of three new species from the Ryukyu Islands: C. (Ceratocombus) ishigaki sp. nov., C. (Xylonannus) meridianus sp. nov. and C. (X.) trimaculatus sp. nov. These new species were assigned to the subgeneric placement, based on the presence of ocular setae, type of vestiture on the pronotum, tarsal formula and venation of the forewing, as noted by previous authors.

Habitus images and line drawings of diagnostic features, including genital structures, are provided. Subgeneric assignments and bionomics were discussed for the three new species. A key to Japanese species of the genus and a checklist of the congeners known in the Palaearctic and Oriental Regions are also provided.