New records of Zachvatkinibates (Acari, Oribatida, Punctoribatidae) in Russia, with description of ontogenetic instars of amphi-Pacific littoral Z. nortoni

Zachvatkinibates nortoni Behan-Pelletier & Eamer, 2005 and Z. erimo Shimano & Aoki, 2019 (Oribatida, Punctoribatidae) are recorded for the first time in Russia, from the seashore of the Magadan Region, North-East Asia. A supplementary description of the adult Z. nortoni and a description of its juvenile instars are presented. Differences between the juveniles of Z. nortoni and those of other Zachvatkinibates species for which juveniles instars have been described are provided.

An identification key and a summary of the distribution and habitats of all Zachvatkinibates known from Russia are presented, along with keys to the known larvae and tritonymphs. The strictly limited littoral habitat preference of Z. nortoni and Z. erimo is accompanied by a highly flexible diet. Of the 19 known species in the genus Zachvatkinibates, nine larger-sized species are associated with seashores, while the remaining ten inhabit various mountain environments, including coniferous forests, steppes, alpine meadows, and barren areas.